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  1. Balsamo,M. & Todaro,M.A. (1987) Aspidiophorus polystictos, a new marine species (Gastrotricha, Chaetonotida) and its life cycle. Boll.Zool. 54, 147-153. Pdf
  2. Balsamo,M. & Todaro,M.A. (1988) Life history traits of two chaetonotids (Gastrotricha) under different experimental conditions. Invert.Reprod.Dev. 14, 161-176. Pdf
  3. Todaro,M.A., Balsamo,M., & Tongiorgi,P. (1988) Tetranchyroderma sardum, a new species of the family Thaumastodermatidae (Gastrotricha, Macrodasyida). Boll.Zool. 55, 69-72. Pdf
  4. Hummon,W.D., Todaro,M.A., Balsamo,M., & Tongiorgi,P. (1990) Effects of pollution on marine Gastrotricha in the northwestern Adriatic Sea. Mar.Pollut.Bull. 21, 241-243. Pdf
  5. Balsamo,M., Todaro,M.A., & Tongiorgi,P. (1992) Marine gastrotrichs from the Tuscan Archipelago (Tyrrhenian Sea): II. Chaetonotida, with description of three new species. Boll.Zool. 59, 487-498. Pdf
  6. Hummon,W.D., Balsamo,M., & Todaro,M.A. (1992) Italian marine Gastrotricha: I. Six new and one redescribed species of Chaetonotida. Boll.Zool. 59, 499-516. Pdf
  7. Todaro,M.A., Balsamo,M., & Tongiorgi,P. (1992) Marine gastrotrichs from the Tuscan Archipelago (Tyrrhenian Sea): I. Macrodasyida, with description of three new species. Boll.Zool. 59, 471-485. Pdf
  8. Todaro,M.A. (1992) Contribution to the study of the Mediterranean meiofauna: Gastrotricha from the Island of Ponza, Italy. Boll.Zool. 59, 321-333. Pdf
  9. Evans,W.A., Todaro,M.A., & Hummon,W.D. (1993) Eutrophication and gastrotrich diversity in the Northern Adriatic Sea. Mar.Pollut.Bull. 26, 268-272. Pdf
  10. Hummon,W.D., Todaro,M.A., & Tongiorgi,P. (1993) Italian Marine Gastrotricha: II. One new genus and ten new species of Macrodasyida. Boll.Zool. 60, 109-127. Pdf
  11. Carman,K.R., Fleeger,J.W., Means,J.C., Pomarico,S.M., McMillin,D.J., Conn,C., & Todaro,M.A. (1994) Experimental investigation of the effects of aromatic hydrocarbons on a sediment food web OCS Study/MMS 94-0033. U.S. Dept. of Interior, Mineral Management Service, Gulf, New Orleans, Louisiana, 89 pp. Pdf
  12. Todaro,M.A. (1994) Chaetonotus triacantus and Heteroxenotrichula texana,two new chaetonotid gastrotrichs from the Gulf of Mexico. Trans.Am.Microsc.Soc. 113, 15-21. Pdf
  13. Balsamo,M. & Todaro,M.A. (1995) Gastrotrichi del Trentino: le Viotte del Monte Bondone (Trento). Studi trent.Sci.nat. 70, 9-22. Pdf
  14. Balsamo,M. & Todaro,M.A. (1995) La meiofauna: una risorsa per l'acquacoltura ed uno strumento per il controllo ambientale. Agribusiness Management & Ambiente 2, 28-42. Pdf
  15. Todaro,M.A. & Balsamo,M. (1995) Marine gastrotrichs from Sicily (Italy). Biol.Gallo-Hellenica 22, 291-292. Pdf
  16. Todaro,M.A. (1995) Paraturbanella solitaria a new psammic species (Gastrotricha: Macrodasyida), from the coast of California. Proc.Biol.Soc.Wash. 108, 553-559. Pdf
  17. Todaro,M.A., Fleeger,J.W., & Hummon,W.D. (1995) Marine gastrotrichs from the sand beaches of the northern Gulf of Mexico: species list and distribution. Hydrobiologia 310, 107-117. Pdf

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  1. Carman,K.R. & Todaro,M.A. (1996) Influence of polycyclic aromatic hydrocarbons on the meiobenthic-copepod community of a Louisiana salt marsh. J.Exp.Mar.Biol.Ecol. 198, 37-54. Pdf
  2. Fleeger,J.W., Shirley,T.C., Carls,M.G., & Todaro,M.A. (1996) Meiofaunal recolonization experiment in oiled sediments. Proceedings of the Exxon Valdez oil spill symposium (ed. by S.Rice, D.A.Spies, D.A.Wolfe, and B.A.Wright), pp. 271-285. American Fisheries Society Symposium, Bethesda,MD. Pdf
  3. Hummon,W.D., Todaro,M.A., Balsamo,M., & Tongiorgi,P. (1996) Italian marine Gastrotricha: III. Four new pentancrous species of the genus Tetranchyroderma (Macrodasyida, Thaumastodermatidae). Ital.J.Zool. 63, 73-79. Pdf
  4. Reible,D.D., Popov,V., Valsaraj,K.T., Thibodeaux,L.J., Lin,F., Dikshit,M., Todaro,M.A., & Fleeger,J.W. (1996) Contaminant fluxes from sediment due to Tubifex bioturbation. Wat.Res. 30, 704-714. Pdf
  5. Todaro,M.A., Fleeger,J.W., Hu,Y.P., Hrincevich,A.W., & Foltz,D.W. (1996) Are meiofauna species cosmopolitan? Morphological and molecular analysis of Xenotrichula intermedia (Gastrotricha: Chaetonotida). Mar.Biol. 125, 735-742. Pdf
  6. Balsamo,M., Hummon,W.D., Todaro,M.A., & Tongiorgi,P. (1997) Italian marine Gastrotricha: IV. Four new species of Chaetonotida. Ital.J.Zool. 64, 83-89. Pdf
  7. Huys,R. & Todaro,M.A. (1997) Meloriastacus ctenidis gen. et sp. nov.: a primitive interstitial copepod (Harpacticoida, Leptastacidae) from Tuscany. Ital.J.Zool. 64, 181-196. Pdf
  8. Hummon,W.D., Todaro,M.A., Tongiorgi,P., & Balsamo,M. (1998) Italian marine Gastrotricha: V. Four new and one redescribed species of Macrodasyida in the Dactylopodolidae and Thaumastodermatidae. Ital.J.Zool. 65, 109-119. Pdf
  9. Todaro,M.A. (1998) Meiofauna from the Meloria Shoals: Gastrotricha, biodiversity and seasonal dynamics. Biol.Mar.Medit. 5, 587-590. Pdf
  10. Todaro,M.A. & Huys,R. (1998) Meiofauna from the Meloria Shoals: observation on a new Leptastacidae (Copepoda, Harpacticoida). Biol.Mar.Medit. 5, 591-595. Pdf
  11. Todaro,M.A. & Kristensen,R.M. (1998) A new species and first report of the genus Nanaloricus (Loricifera, Nanaloricida, Nanaloaricidae) from the Mediterranean Sea. Ital.J.Zool. 65, 219-226. Pdf
  12. Todaro,M.A. (1998) Meiofauna, non solo look. Naturalmente 11, 43-47. Pdf-->
  13. Todaro,M.A. (1998) Note faunistiche ed ecologiche sui Gastrotrichi delle Secche della Meloria. Fauna nel Mediteraneo, immagini e note di ecologia marina. Quaderni dell'Acquario (Livorno) pp. 57-72. Quaderni dell'Acquario, vol. 3, Livorno. Pdf
  14. Todaro, M. A. La meiofauna del sedimento organogeno grossolano delle Secche della Meloria: biodiversità ed analisi morfologiche e molecolari per riscontri filogenetici e biogeografici. 1-146. 1998. Universita di Bologna. 1-1-1998. Ref Type: Thesis/Dissertation Pdf-->
  15. Wilson,C.A., Nieland,D.L., Fleeger,J.W., Todaro,M.A., Malone,R.F., & Rusch,K.A. (1998) Preliminary evaluation of the use of phosphogypsum for reef substrate. II. A study of the effects of phosphogypsum exposure on diversity and biomass of acquatic organisms. Chem.Ecol. 14, 321-340. Pdf

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  1. D' Addabbo,M., Grimaldi-De Zio,S., Morone,M.R.D., Pietanza,R., D' Addabbo,R., & Todaro,M.A. (1999) Diversity and dynamics of an interstitial Tardigrada population in the Meloria Shoals, Ligurian Sea, with a redescription of Batillipes similis (Heterotardigrada, Batillipedidae). Ital.J.Zool. 66, 51-61. Pdf
  2. Todaro,M.A. (1999) Copepodi arpacticoidi delle secche della Meloria: segnalazione di generi nuovi per l'Italia. Biologia Marina Mediterranea 6, 452-456. Pdf
  3. Casu,D., Milella,I., Castelli,A., & Todaro,M.A. (2000) Meiobenthic polychaetes from the Meloria Shoals. Biol.Mar.Medit. 7, 666-669. Pdf
  4. Todaro,M.A., Bernhard,J.M., & Hummon,W.D. (2000) A new species of Urodasys (Gastrotricha, Macrodasyida) from dysoxic sediments of the Santa Barbara Basin (California, USA). Bull.Mar.Sci. 66, 467-476. Pdf
  5. Todaro,M.A., Virno-Lamberti,C., Pulcini,M., Pellegrini,D., & De Ranieri,S. (2001) Monitoring of a dredged material disposal site: evidence of rapid meiofaunal recolonization of the dumped sediments. Biol.Mar.Medit. 8, 1-10. Pdf
  6. Todaro,M.A., Faranapova,O., Onorati,F., Pellegrini,D., & Tongiorgi,P. (2001) Tigripus fulvus (Copepoda, Harpacticoida) a possible test-species in harbour sediment toxicity bioassessment: life cycle and preliminary bioassays. Biol.Mar.Medit. 8, 1-10. Pdf
  7. Todaro,M.A., Hummon,W.D., Balsamo,M., Fregni,E., & Tongiorgi,P. (2001) Inventario dei Gastrotrichi marini italiani: una checklist annotata. Atti Soc.Tosc.Sci.Nat.Mem.Serie B 107, 75-137. Pdf
  8. Balsamo,M., Ferraguti,M., Guidi,L., Todaro,M.A., & Tongiorgi,P. (2002) Reproductive system and spermatozoa of Paraturbanella teissieri (Gastrotricha, Macrodasyida): implications for sperm transfer modality in Turbanellidae. Zoomorphology 121, 235-241. Pdf
  9. Balsamo,M. & Todaro,M.A. (2002) Gastrotricha. Freshwater Meiofauna Biology and Ecology (ed. by S.D.Rundle, A.I.Robertson, and J.M.Schmid-Araya), pp. 45-61. Backhuys Publisher, Leiden. Pdf
  10. Giudici,P., Pulvirenti,A., Todaro,M.A., & Zambonelli,C. (2002) Horizontal transfer of genetic materials in Saccharomyces promoted by animals. Bulletin de Office international de la vigne et du vin (l'O.I.V.) 75, 484-500. Pdf
  11. Pulvirenti,A., Zambonelli,C., Todaro,M.A., & Giudici,P. (2002) Interspecific hybridation by digestive tract of invertebrates as sorce of environmental biodiversity within the Saccaromyces cerevisiae. Ann.Microbiol. 52 , 245-255. Pdf
  12. Todaro,M.A., Balsamo,M., & Tongiorgi,P. (2002) Marine gastrotrich fauna in Corsica (France), with a description of a new species of the genus Tetranchyroderma (Macrodasyida, Thaumastodermatidae). Sarsia 87, 248-257. Pdf
  13. Todaro,M.A. (2002) An interesting new gastrotrich from littoral meiobenthos (Long Beach Island, USA), with a key to species of Tetranchyroderma (Gastrotricha : Macrodasyida). J.Mar.Biol.Assoc.U.K. 82, 555-563. Pdf
  14. Todaro,M.A., Virno-Lamberti,C., Pulcini,M., Pellegrini,D., & De Ranieri,S. (2002) Meiofauna of a dredging disposal site. Biol.Mar.Medit. 9, 647-650. Pdf
  15. Baraldi,E., Virno-Lamberti,C., Pellegrini,D., Leasi,F., & Todaro,M.A. (2003) Biomonitoring of a dumping area: the meiobenthic community. Biol.Mar.Medit. 10, 640-643. Pdf

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  1. Faranapova,O., Todaro,M.A., Onorati,F., & Finoia,M.G. (2003) Sex and age sensitivity of Tigriopus fulvus (Copepoda, Harpacticoida) towards two heavy metals (Cd and Cu). Biol.Mar.Medit. 10, 676-681. Pdf
  2. Guidi,L., Marotta,R., Pierboni,L., Ferraguti,M., Todaro,M.A., & Balsamo,M. (2003) Comparative sperm ultrastructure of Neodasys ciritus and Musellifer delamarei, two species considered to be basal among Chaetonotida (Gastrotricha). Zoomorphology 122, 135-143. Pdf
  3. Todaro,M.A., Matinato,L., Balsamo,M., & Tongiorgi,P. (2003) Faunistics and zoogeographical overview of the Mediterranean and Black Sea marine Gastrotricha. Biogeographia 24, 131-160. Pdf
  4. Todaro,M.A. & Shirley,T.C. (2003) A new meiobenthic priapulid (Priapulida, Tubiluchidae) from a Mediterranean submarine cave. Ital.J.Zool. 70, 79-87. Pdf
  5. Todaro,M.A., Ancona,P., Marzano,A., D' Addabbo,M.G., & De Zio-Grimaldi,S. (2003) A new Tetranchyroderma species (Gastrotricha, Macrodasyida, Thaumastodermatidae) from the Canary Islands (Spain). Cah.Biol.Mar. 44, 191-197. Pdf
  6. Todaro,M.A., Littlewood,D.T.J., Balsamo,M., Herniou,E.A., Cassanelli,S., Manicardi,G., Wirz,A., & Tongiorgi,P. (2003) The interrelationships of the Gastrotricha using nuclear small rRNA subunit sequence data, with an interpretation based on morphology. Zool. Anz. 242, 145-156. Pdf
  7. Ansaloni,I., Baraldi,E., Mauri,M., Montanari,G., Pagliai,A.M., Preti,M., Prevedelli,D., Rinaldi,A., Simonini,R., & Todaro,M.A. (2004) Effetti dell'estrazione di sabbie marine sulla comunità macrozoobentonica delle sabbie relitte dell'Adriatico settentrionale. In: Casagrandi, R. & Melià, P. (Eds.) Ecologia. Atti XIII Congresso SITe. Pdf
  8. Balsamo,M., Todaro,M.A., & Tongiorgi,P. (2004) Gastrotricha. Freshwater Invertebrates of the Malaysian regio (ed. by C.M.Yule and H.S.Yong), pp. 127-135. Academy of Science Malaysia. Pdf
  9. Guidi,L., Pierboni,L., Ferraguti,M., Todaro,M.A., & Balsamo,M. (2004) Spermatology of the genus Lepidodasys Remane, 1926 (Gastrotricha, Macrodasyida): towards a revision of the family Lepidodasyidae Remane, 1927. Acta Zool. 85, 211-221. Pdf
  10. Todaro,M.A. & Rocha,C.E.F. (2004) Diversity and distribution of marine Gastrotricha along the northern beaches of the state of Sao Paulo (Brazil), with description of a new species of Macrodasys (Macrodasyida, Macrodasyidae). J.Nat.Hist. 38, 1605-1634. Pdf
  11. Hummon,W.D., Todaro,M.A., & Evans,W.A. (2005) Video database for described species of marine Gastrotricha. Meiofauna Mar. 14, 23-26. Pdf
  12. Marotta,R., Guidi,L., Pierboni,L., Ferraguti,M., Todaro,M.A., & Balsamo,M. (2005) Sperm ultrastructure of Macrodasys caudatus (Gastrotricha: Macrodasyida) and a sperm based phylogenetic analysis of Gastrotricha. Meiofauna Mar. 14, 9-21. Pdf
  13. Todaro,M.A. & Rocha,C.E.F. (2005) Further data on marine gastrotrichs from the State of São Paulo and the first records from the State of Rio de Janeiro (Brazil). Meiofauna Mar. 14, 27-31. Pdf
  14. Todaro,M.A., Balsamo,M., & Kristensen,R.M. (2005) A new genus of marine chaetonotids (Gastrotricha), with a description of two new species from Greenland and Denmark. J.Mar.Biol.Assoc.U.K. 85, 1391-1400. Pdf

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  1. Nicholas,N. & Todaro,M.A. (2005) Observations on Gastrotricha from a sandy beach in southeastern Australia with a description of Halichaetonotus australis sp. nov. (Gastrotricha: Chaetonotida). N Z.J.Mar Freshwat.Res 39, 973-980. Pdf
  2. Ferrari,G., Ansaloni,I., Mauri,M., Pagliai,A.M., Prevedelli,D., Simonini,R., & Todaro,M.A. (2006) La meiofauna delle sabbie 'relitte' dell'Adriatico settentrionale soggette ad operazioni di escavo. Biol.Mar.Medit. 13, 574-578. Pdf
  3. Leasi,F., Virno-Lamberti,C., & Todaro,M.A. (2006) First record of Musellifer profundus (Gastrotricha) in the Italian seas. Biol.Mar.Medit. 13, 190-191. Pdf
  4. Leasi,F., Rothe,B.H., Schmidt-Rhaesa A., & Todaro,M.A. (2006) The musculature of three species of gastrotrichs surveyed with confocal laser scanning microscopy (CLSM). Acta Zool. 87, 171-180. Pdf
  5. Nicholas,W. & Todaro,M.A. (2006) Two new species of Tetranchyroderma (Gastrotricha, Macrodasyida) from a sandy beach in southeastern Australia. N.Z.J.Mar.Freshwat.Res. 40, 249-258. Pdf
  6. Todaro,M.A., Balsamo,M., & Tongiorgi,P. (2006) Gastrotricha. La checklist della fauna marina italiana (ed. by G.Relini and S.La Posta), Ministero dell'Ambiente e della Tutela del territorio. Link
  7. Todaro,M.A. & Leasi,F. (2006) Further data on the Italian marine gastrotrich fauna. Biol.Mar.Medit. 13, 1116-1120. Pdf
  8. Todaro,M.A., Leasi,F., Bizzarri,N., & Tongiorgi,P. (2006) Meiofauna densities and gastrotrich community composition in a Mediterranean sea cave. Mar.Biol. 149, 1079-1091. Pdf
  9. Todaro,M.A. (2006) Loricifera. In: La checklist della fauna marina italiana (ed. by G.Relini and S.La Posta), Ministero dell'Ambiente e della Tutela del territorio. Link
  10. Todaro,M.A. (2006) Priapulida. In: La checklist della fauna marina italiana (ed. by G.Relini and S.La Posta), Ministero dell'Ambiente e della Tutela del territorio. Link
  11. Todaro,M.A. & Ceccherelli,V.U. (2006) Harpacticoida. In: La checklist della fauna marina italiana (ed. by G.Relini and S.La Posta), Ministero dell'Ambiente e della Tutela del territorio. Link
  12. Todaro,M.A., Telford,M.J., Lockyer,A.E., & Littlewood,D.T.J. (2006) Interrelationships of the Gastrotricha and their place among the Metazoa inferred from 18S rRNA genes. Zool.Scr. 35, 251-259. Pdf
  13. Todaro,M.A., Guidi,L., Leasi,F., Tongiorgi,P. (2006) Morphology of Xenodasys (Gastrotricha): the first species from the Mediterranean Sea and the establishment of Chordodasiopsis gen. nov. and Xenodasyidae fam. nov. J.Mar.Biol.Assoc.U.K., 86, 1005-1015. Pdf
  14. Rothe B. H., Schmidt-Rhaesa A., Todaro,M.A. (2006) The general muscular architecture in Tubiluchus troglodytes (Priapulida). Meiofauna Marina (München ) 15: 79-86. Pdf
  15. Balsamo, M., Guidi, L., Pierboni, L., Marotta, R., Todaro, M. A. & Ferraguti, M. 2007. Living without mitochondria : spermatozoa and spermatogenesis in two species of Urodasys (Gastrotricha, Macrodasyida) from dysoxic sediments. Invertebrate Biology 126: 1-9. Pdf
  16. Hummon, W. D. & Todaro, M. A. 2007. A new species of Xenotrichulidae (Gastrotricha) from southern and southeastern USA. Cahiers De Biologie Marine 48: 297-302. Pdf
  17. Priyalakshmi, G., Menon, N. R. & Todaro, M. A. 2007. A new species of Pseudostomella (Gastrotricha: Macrodasyida:Thaumastodermatidae) from a sandy beach of Kerala, India. Zootaxa 1616 : 61-68. Pdf

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  1. Marotta, R., Todaro M.A. & Ferraguti, M. (2008). The unique gravireceptor organs of Pleurodasys helgolandicus (Gastrotricha: Macrodasyida). Zoomorphology 127: 111-119. Pdf
  2. Leasi, F. & Todaro, M. A. (2008). The muscular system of Musellifer delamarei (Renaud-Mornant, 1968) and other chaetonotidans with implications for the phylogeny and systematization of the Paucitubulatina (Gastrotricha). Biological Journal of the Linnean Society 94: 379.-398. Pdf
  3. Todaro,M.A. & Hummon, W.D. (2008). An overview and a dichotomous key to genera of the phylum Gastrotricha. Meiofauna Marina 16: 3-20. Pdf
  4. Guidi, L., Ferraguti, M., Todaro, M.A., Pierboni, L., Balsamo, M. (2009). Unusual spermatozoa and reproductive modalities of Xenodasys eknomios (Gastrotricha: Xenodasyidae). Ital. J. Zool. 76: 165–172 Pdf
  5. Leasi, F. & Todaro, M. A. (2009). Meiofaunal cryptic species revealed by confocal microscopy: the case of Xenotrichula intermedia (Gastrotricha). Mar. Biol. 156: 1335-1346. Pdf
  6. Todaro, M. A., Dal Zotto, M., Maiorova, A. S. & Adrianov, A.V. (2009). A new species of Aspidiophorus (Gastrotricha, Chaetonotida) from the Russian Far East with a key to marine species of the genus. Mar.Biol.Res. 5: 297-333. Pdf
  7. Willems, W. R., Curini-Galletti, M., Ferrero, T.J., Fontaneto, D., Heiner, I., Huys, R., Ivanenko, V.N., Kristensen, R.M., Kanneby, T., MacNaughton, M.O., Martínez Arbizu, P., Todaro, M.A., Sterrer W. & Jondelius, U. (2009). Meiofauna of the Koster-area, results from a workshop at the Sven Lovén Centre for Marine Sciences (Tjärnö, Sweden). Meiofauna Marina (München) 17:1-34. Pdf
  8. Dal Zotto, M. & Todaro, M.A. (2009 for 2008). Cycliophora. In: La checklist della flora e della fauna dei mari italiani (a cura di G. Relini). Biol. Mar. Mediterr. 15 (suppl.): 178-181. Pdf
  9. Todaro, M.A. (2009 for 2008) Loricifera. In: La checklist della della flora e della fauna dei mari italiani (a cura di G. Relini). Biol. Mar. Mediterr. 15 (suppl.): 170-173. Pdf
  10. Todaro, M.A. (2009 for 2008) Priapulida. In. La checklist della flora e della fauna dei mari italiani (a cura di G. Relini). Biol. Mar. Mediterr. 15 (suppl.): 174-177. Pdf
  11. Todaro, M.A., Balsamo, M. & Tongiorgi, P. (2009 for 2008) Gastrotricha. In: La checklist della flora e della fauna dei mari italiani (a cura di G. Relini). Biol. Mar. Mediterr., 15 (suppl.): 160-169. Pdf
  12. Kanneby, T., Todaro, M. A. & Jondelius, U. (2009) One new species and records of Ichthydium Ehrenberg, 1830 (Gastrotricha: Chaetonotida) from Sweden with a key to the genus. Zootaxa 2278: 26-46. Pdf
  13. Orsi, C. F., Colombari, B., Callegari, F., Todaro, M. A., Ardizzoni, A., Rossini, G. P., Blasi, E. & Peppoloni, S. (2009). Yessotoxin inhibits phagocytic activity of macrophages. Toxicon 55: 265-273. Pdf
  14. Hummon, W. D. & Todaro, M. A. (2009) Italian marine Gastrotricha: VI. Seven new species of Macrodasyida. Zootaxa 2278: 47-68. Pdf
  15. Dal Zotto, M. & Todaro, M. A. (2009) Una nuova species di Antigomonas (Kinorhyncha: Cyclorhagida) dalla costa toscana. Biol. Mar. Medit. 16: 258-259. Pdf

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  1. Hummon, W.D. & Todaro, M.A. (2010) Analytic taxonomy and notes on marine, brackish-water and estuarine Gastrotricha. Zootaxa 2392: 1–32. Pdf
  2. Leasi, F. & Todaro, M.A. (2010) The gastrotrich community of a north Adriatic Sea site, with a redescription of Musellifer profundus (Chaetonotida: Muselliferidae). J. Mar. Biol. Ass. U.K. 90: 645-653. doi:10.1017/S0025315409991068. Pdf
  3. Todaro, M.A. & Checcherelli, V.U. (2010) Harpacticoida. In checklist della Flora e della Fauna delle specie marine italiane (parte II). Biol. Mar. Mediter. 17 (suppl.): 452-464. Pdf
  4. Balsamo, M., Albertelli, G., Ceccherelli, V. U., Coccioni, R., Colangelo, M. A., Curini-Galletti, M., Danovaro, R., D'Addabbo, R., De Leonardis, C., Fabiano, M., Frontalini, F., Gallo, M., Gambi, C., Guidi, L., Moreno, M., Pusceddu, A., Sandulli, R., Semprucci, F., Todaro, M. A. & Tongiorgi, P. 2010. Meiofauna of the Adriatic Sea: present knowledge and future perspectives. Chemistry and Ecology 26: 45-63. Pdf
  5. Dal Zotto, M., Ghiviriga, S. & Todaro, M.A. 2010. A new Tetranchyroderma (Gastrotricha, Thaumastodermatidae) with triancres from the Mediterranean Sea. Meiofauna Marina 18: 41-48. Pdf
  6. Todaro, M.A., Kanneby, T., Dal Zotto, M. & Jondelius, U. 2011. Phylogeny of Thaumastodermatidae (Gastrotricha: Macrodasyida) Inferred from Nuclear and Mitochondrial Sequence Data. PLoS ONE 6: e17892. Pdf
  7. Guidi, L., Todaro, M. A., Ferraguti, M. & Balsamo, M. 2011. Reproductive system of the genus Crasiella (Gastrotricha, Macrodasyida). Helgoland Marine Research 65: 175-185. Pdf
  8. Artois, T., Fontaneto, D., Hummon, W. D., Mcinnes, S. J., Todaro, M. A., Sorensen, M. V. & Zullini, A. 2011. Ubiquity of microscopic animals? Evidence from the morphological approach in species identification. In Fontaneto, D. (Ed) Biogeography of microscopic organisms: is everything small everywhere? (pp. 244-283). Cambridge: Cambridge University Press. Pdf
  9. Todaro, M. A., Dal Zotto, M., Perissinotto, R. & Bownes, S. J. 2011. First records of Gastrotricha from South Africa, with description of a new species of Halichaetonotus (Chaetonotida, Chaetonotidae). Zookeys 142: 1-13. Pdf
  10. Marotta, R., Balsamo, M., Ferraguti, M., Fondello, C., Guidi, L. & Todaro, M. A. 2011. The spermatozoon of Thaumastoderma moebjergi with the description of the sperm model for the family Thaumastodermatidae (Gastrotricha, Macrodasyida). Meiofauna Marina 19: 89-95. Pdf
  11. Rocha, C. E. F., Kihara, T. C., Todaro, M. A. , Hooge, M. & Jonnson, R. 2011. Meiofauna. In Amaral, A. C. Z. & Nallin, S. A. H. (Eds) Biodiversidade e ecossistemas bentônicos marinhos do Litoral Norte de São Paulo, Sudeste do Brasil. (pp. 43-47). Campinas: Campinas, SP:IB/UNICAMP. Pdf
  12. Todaro, M. A. & Rocha, C. E. F. 2011. Gastrotricha. In Amaral, A. C. Z. & Nallin, S. A. H. (Eds) Biodiversidade e ecossistemas bentônicos marinhos do Litoral Norte de São Paulo, Sudeste do Brasil (pp. 100-105). Campinas: Campinas, SP: IB/UNICAMP. Pdf

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  1. Kanneby T., Todaro M. A. & Jondelius U. 2012. A phylogenetic approach to species delimitation in freshwater Gastrotricha from Sweden. Hydrobiologia. 683: 185-202. DOI 10.1007/s10750-011-0956-1 Pdf
  2. Todaro M.A., Dal Zotto M., Jondelius U., Hochberg R., Hummon W.D., Kanneby T., Rocha C.E.F. 2012. Gastrotricha: A Marine Sister for a Freshwater Puzzle. PLoS ONE 7(2): e31740. doi:10.1371/journal.pone.0031740 Pdf
  3. Daly M., Perissinotto R., Laird M., Dyer D., Todaro M. A. 2012. Description and ecology of a new species of Edwardsia de Quatrefages, 1842 (Anthozoa, Actiniaria) from the St Lucia Estuary, South Africa. Marine Biology Research 8: 233-245. Pdf
  4. Curini-Galletti M., Artois T., Delogu V., De Smet W.H., Fontaneto D., Jondelius U., Leasi F., Martinez A., Meyer-Wachsmuth I., Nilsson K.S., Tongiorgi P., Worsaae K. & Todaro M.A. 2012. Patterns of Diversity in Soft-Bodied Meiofauna: Dispersal Ability and Body Size Matter. PLoS ONE 7(3): e33801. doi:10.1371/journal.pone.0033801. Pdf
  5. Todaro M.A., Guid L., Ferraguti M. & Balsamo M. 2012. A fresh look at Dinodasys mirabilis (Gastrotricha, Macrodasyida), with focus on the reproductive apparatus and sperm ultrastructure. Zoomorphology 131: 115-125. Pdf
  6. Kanneby, T., Todaro, M. A. & Jondelius, U. 2012. Phylogeny of Chaetonotidae and other Paucitubulatina (Gastrotricha: Chaetonotida) and the colonization of aquatic ecosystems. Zoologica Scripta doi:10.1111/j.1463-6409.2012.00558.x. Pdf
  7. Todaro M.A. 2012. A new marine gastrotrich from the State of São Paulo (Brazil), with a key to species of Pseudostomella (Gastrotricha, Thaumastodermatidae). ZooKeys 223: 39–51. Pdf
  8. Riera R., Todaro M. A. 2012. Check list of gastrotrichs from the Canary Islands. Rev.Acad.Canar.Cienc. 24: 81-88. Pdf
  9. Appeltans W., Ahyong S., ........, Todaro M.A. et al. 2012. The Magnitude of Global Marine Species Diversity. Current Biology 22: 2189–2202. Pdf
  10. Todaro M.A., Leasi F. 2013. A new eye-bearing Macrodasys (Gastrotricha : Macrodasyida) from Jamaica. Meiofauna Marina 20: 33-38. Pdf
  11. Todaro M. A. 2013. A new non-naked species of Ptychostomella (Gastrotricha) from Brazil. ZooKeys 289: 13–24. Pdf
  12. Todaro M.A., Perissinotto R., Bownes S. J. 2013. Neogosseidae (Gastrotricha, Chaetonotida) from the iSimangaliso Wetland Park, KwaZulu-Natal, South Africa. Zookeys 315: 77-94. Pdf
  13. Guidi, L., Todaro, M. A., Ferraguti, M. & Balsamo, M. 2014. Reproductive system and spermatozoa ultrastructure support the phylogenetic proximity of Megadasys and Crasiella (Gastrotricha, Macrodasyida). Contributions to Zoology 83: 119-131. Pdf
  14. Todaro, M.A., Leasi, F. & Hochberg, R. 2014. A new species, genus and family of marine Gastrotricha from Jamaica, with a phylogenetic analysis of Macrodasyida based on molecular data. Systematics and Biodiversity. 12: 473-488. Pdf

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  1. Todaro M. A., Perissinotto R. & Bownes S.J. 2015.Two new marine Gastrotricha from the Indian Ocean coast of South Africa. Zootaxa 3905 (2): 193–208. http://dx.doi.org/10.11646/zootaxa.3905.2.2. Pdf
  2. Egger B., Lapraz F., Tomiczek B., Müller S., Dessimoz C., Girstmair J., Skunca N., Rawlinson K.A., Cameron C.B., Beli E., Todaro M.A., Gammoudi M., Noreña C. & Telford M.J. 2015. A Transcriptomic-Phylogenomic Analysis of the Evolutionary Relationships of Flatworms. Current Biology 25: 1–7,http://dx.doi.org/10.1016/j.cub.2015.03.034 Pdf
  3. Kånneby T. & Todaro M.A. 2015. The phylogenetic position of Neogosseidae (Gastrotricha: Chaetonotida) and the origin of planktonic Gastrotricha. Organisms Diversity & Evolution. 15: 459-469. DOI.10.1007/s13127-015-0223-9 Pdf
  4. Todaro M.A., Dal Zotto M. & Leasi F. 2015. An integrated morphological and molecular approach to the description and systematisation of a novel genus and species of Macrodasyida (Gastrotricha). PLoS ONE 10(7): e0130278. doi:10.1371/journal.pone.0130278 Pdf
  5. Balsamo M., d`Hondt J-L., Kisielewski J., Todaro M.A., Tongiorgi P., Guidi L., de Jong Y. 2015. Fauna Europaea: Gastrotricha. Biodiversity Data Journal 3: e5800. doi:10.3897/BDJ.3.e5800 Pdf
  6. Bernhard J. M., Morrison C. R., Pape E., Beaudoin D. J., Todaro M. A., Pachiadaki M. G., Kormas K. A. & Edgcomb V. P. 2015. Metazoans of redoxcline sediments in Mediterranean deep-sea hypersaline anoxic basins. BMC Biology 13:105. doi:10.1186/s12915-015-0213-6 Pdf
  7. Dal Zotto M., Todaro M.A. 2016. Kinorhyncha from Italy, a revision of the current checklist and an account of the recent investigations. Zoologisher Anzeiger. Doi: 10.1016/j.jcz.2016.01.004 Pdf
  8. Dal Zotto M., Santulli A., Simonini R., Todaro M.A. 2016. Organic enrichment effects on a marine meiofauna community, with focus on Kinorhyncha. Zoologisher Anzeiger. Doi: 10.1016/j.jcz.2016.03.013 Pdf
  9. Baucon A., de Carvalho C.N., Barbieri R., Bernardini F., Cavalazzi B., Celani A., Felletti F., Ferretti A., Schönlaub H.P., Todaro A., Tuniz C. 2017. Organism-substrate interactions and astrobiology: Potential, models and Methods. Earth-Science Review 171:141-180. Pdf
  10. Guidi L., Todaro M.A., Cesaroni L., Balsamo M. 2017. The unusual spermatozoa of Dolichodasys sp. (Gastrotricha, Macrodasyida). Proceedings of the Biological society of Washington 130:186–196. Pdf
  11. Kånneby T., Todaro M. A. 2017. A new species of Aspidiophorus (Gastrotricha: Chaetonotidae) from the Swedish west coast Zootaxa 4290: 390-394. Pdf
  12. Todaro M.A., Dal Zotto M., Bownes S.J., Perissinotto R. 2017. Two new interesting species of Macrodasyida (Gastrotricha) from KwaZulu-Natal (South Africa). Proceedings of the Biological society of Washington 130:139–154. Pdf
  13. Todaro M.A., Dal Zotto M., Cesaretti. A. 2017. Marine gastrotrichs from Lanzarote, w ith a description of a phylogenetically relev ant species of Urodasys (Gastrotricha, Macrodasyida). Marine Biodiversity. doi:10.1007/s12526-017-0747-7 Pdf

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  1. Dal Zotto M., Leasi F., Todaro M.A. 2018. A new species of Turbanellidae (Gastrotricha, Macrodasyida) from Jamaica, with a key to species of Paraturbanella. Zookeys 734: 105-119. doi: 10.3897/zookeys.734.23023 Pdf
  2. Schuster J., Atherton S., Todaro M. A., Schmidt-Rhaesa A., Hochberg R. 2018. Redescription of Xenodasys riedli (Gastrotricha: Macrodasyida) based on SEM analysis, with first report of population density data. Marine Biodiversity 48: 259-271. DOI 10.1007/s12526-017-0667-617 Pdf
  3. Leasi F., Joseph L. Sevigny J.L., , Eric M. Laflamme E.M., Artois T., Curini-Galletti M., de Jesus Navarrete A., Di Domenico M., Goetz F., Hall J.A., Hochberg R., Jörger K.M., Jondelius U., Todaro M.A., Wirshing H.H., Jon L. Norenburg J.L., Thomas W. K. 2018. Biodiversity estimates and ecological interpretations of meiofaunal communities are biased by the taxonomic approach. Communication Biology 1: 112. DOI: 10.1038/s42003-018-0119-2 Pdf
  4. Chatterjee T., Priyalakshmi G., Todaro M.A. 2019. An annotated checklist of the macrodasyidan Gastrotricha from India. Zootaxa 4545 (4): 495–510. Pdf
  5. Sergeeva N.G., Ürkmez D., Todaro M.A. 2019. Significant occurrence of Musellifer profundus Vivier, 1974 (Gastrotricha, Chaetonotida) in the Black Sea. Check List 15 (1): 219–224. Pdf
  6. Schwaha T., Bernhard J.M., Edgcomb V.P., Todaro M.A. 2019. Aethozooides uraniae, a new deep-sea genus and species of solitary bryozoan from the Mediterranean Sea, with a revision of the Aethozoidae. Marine Biodiversity. https://doi.org/10.1007/s12526-019-00948-w Pdf
  7. Todaro M.A., Dal Zotto M., Kånneby T., Hochberg R. 2019. Integrated data analysis allows the establishment of a new, cosmopolitan genus of marine Macrodasyida (Gastrotricha). Scientific Reports 9:7989. https://doi.org/10.1038/s41598-019-43977-y Pdf
  8. Todaro M.A., Sibaja-Cordero J.A., O.A.,Coto-Delgado G., Goebel-Otárola N., Barquero J.D., Mariana Cullell-Delgado M., Dal Zotto M. 2019. An introduction to the study of Gastrotricha, with a taxonomic key to families and genera of the group. Diversity 11: 117. doi:10.3390/d11070117 Pdf
  9. Dal Zotto M., Neuhaus B., Yamasaki H., Todaro M.A. 2019. The genus Condyloderes (Kinorhyncha: Cyclorhagida) in the Mediterranean Sea, including the description of two new species with novel characters. Zoologischer Anzeiger 282: 206-231 Pdf
  10. Martínez A., Di Domenico M., Leasi F., Curini-Galletti M., Todaro M.A., Dal Zotto M., Gobert S., Artois T., Norenburg J., Jörger K.M., Núñez J., Fontaneto D., Worsaae K. 2019. Patterns of diversity and endemism of soft-bodied meiofauna in an oceanic island, Lanzarote, Canary Islands. Marine Biodiversity doi: 10.1007/s12526-019-01007-0 Pdf

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  1. Martínez A., Eckert, E. M., Artois T., Careddu G., Casu M., Curini-Galletti M., Gazale V., Gobert S., Ivanenko V.N., Jondelius U., Marzano M., Pesole G., Zanello A., Todaro M.A., Fontaneto D. 2020. Human access impacts biodiversity of microscopic animals in sandy beaches. Communications Biology 3:175. https://doi.org/10.1038/s42003-020-0912-6 Pdf
  2. Campos A., Todaro, M.A., Garraffoni A.R.S. 2020. A New Species of Paraturbanella Remane, 1927 (Gastrotricha, Macrodasyida) from the Brazilian Coast, and the Molecular Phylogeny of Turbanellidae Remane, 1926. Diversity-Basel 12. Pdf
  3. Balsamo M., Artois, T., Smith, J. P. S., Todaro, M. A., Guidi, L., Leander, B. S. & Van Steenkiste, N. W. L. 2020. The curious and neglected soft-bodied meiofauna: Rouphozoa (Gastrotricha and Platyhelminthes). Hydrobiologia 847: 2613-2644. Pdf
  4. Curini-Galletti, M., Artois, T., Di Domenico, M., Fontaneto, D., Jondelius, U., Jorger, K. M., Leasi, F., Martinez, A., Norenburg, J. L., Sterrer, W. & Todaro, M. A. 2020. Contribution of soft-bodied meiofaunal taxa to Italian marine biodiversity. European Zoological Journal 87: 369-384. Pdf
  5. Dal Zotto M., De Biasi A.M., Pacciardi L., Pertusati M., Bianchi V., Giannetti M., Rebecchi C. & Todaro M.A. 2020. Studio della meiofauna nei pressi di un rigassificatore offshore. Biologia Marina Mediterranea 26: 103-104. Pdf
  6. Todaro M.A., Dal Zotto M., Rebecchi C. 2020. Presenza del raro Maccabeus tentaculatus Por, 1973 (Priapulida) nel Mar Ligure. Biologia Marina Mediterranea 26: 326-327. Pdf
  7. Chatterjee T., Todaro M.A. 2021. An annotated checklist of the Chaetonotidan Gastrotricha from India. Zootaxa 4545: 495-510. Pdf
  8. Kieneke, A. & Todaro, M. A. 2021. Discovery of two 'chimeric' Gastrotricha and their systematic placement based on an integrative approach. Zoological Journal of the Linnean Society 192: 710-735. Pdf
  9. Guidi L., Balsamo M., Ferraguti M., Todaro M. A. 2021. Reproductive organs and spermatogenesis of the peculiar spermatozoa of the genus Kryptodasys (Gastrotricha, Macrodasyida), with an appraisal of the occurrence and origin of the tail-less spermatozoa in Gastrotricha. J. Zool. Syst. Evol. Res. DOI: 10.1111/jzs.12556 Pdf
  10. Todaro M.A., Balsamo M., 2021. Gastrotricha. In: Bologna M.A., Zapparoli M., Oliverio M., Minelli A., Bonato L., Cianferoni F., Stoch F. (eds.), Checklist of the Italian Fauna. Version 1.0. Last update: 2021-05-31. Available at: https://www.lifewatchitaly.eu/en/initiatives/checklist-fauna-italia-en/checklist/
  11. Todaro M.A., Luporini P. 2022. Not too big for its mouth: direct evidence of a macrodasyidan gastrotrich preyed in nature by a dileptid ciliate. The European Zoological Journal, 89:785-790. DOI: 10.1080/24750263.2022.2095048 Pdf

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  1. Cesaretti A., F. Leasi F., M.A. Todaro M.A. 2023. Confocal Laser Scanning Microscopy Applied to a New Species Helps Understand the Functioning of the Reproductive Apparatus in Stylet-Bearing Urodasys (Gastrotricha: Macrodasyida). Water 15: 1106; https://doi.org/10.3390/w15061106 Pdf
  2. Todaro M.A., Anselmi S., Bentivoglio T., Pretti C., Cavallo A., Renzi M. 2023. Looking for Nano- and Microplastics in Meiofauna Using Advanced Methodologies. Environments 2023 10(5):81. https://doi.org/10.3390/environments10050081 Pdf
  3. Todaro, M.A. (2023) Chapter 10. Kingdom Animalia, phylum Gastrotricha (hairy-bellied worms). Pp. 187–191 in: Kelly, M., Mills, S., Terezow, M., Sim-Smith, C., Nelson, W. (Eds) The Marine Biota of Aotearoa New Zealand. Updating our marine biodiversity inventory. NIWA Biodiversity Memoir 136, 494 pp. Pdf
  4. Saponi F., Todaro M.A. 2024. Status of the Italian Freshwater Gastrotricha Biodiversity, with the Creation of an Interactive GIS-Based Web Map. Diversity 16: 17. https://doi.org/10.3390/d16010017 Pdf

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